[MapProxy] seed only question

[DeDuikertjes]

Dear List,

Please help me with the following question:

I have a layer with a cache as source, this cache has a source as source 
with seed_only set to true.
If I request a tile from the layer, the tile is NOT in the cache will 
Mapproxy return a transparent image or a 404-error?

Sam example taken further:
To mapproxy I issue a WMS-request which combines multiple layers, each 
layer has a separate cache, each of these caches a separate read-only 
source. One of the requested tiles is not in cache; what will mapproxy 
return?

thank you very much, MArco
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